want to do that.Torque in gear is the way a car operates.Starting in fifth gear and keeping it there would kill your MPG not help it.Ideally you want it to shift at the torque peak in each gear allowing ease of movement in that gear to the next and so on.Ever drive a stick manual transmission?Try feathering the clutch and taking off in high gear,unless the car has gobs of torque you will fry the clutch,or stall it out,or the engine will lug along straining to come to the torque peak for that gear.Not conducive to high MPG.
Every automatic has a compromise to obtain comfort in shift quality.If you want to maximize MPG get a G force meter and shift in manual mode at peak torque in each gear just short of the highest attained mpg using your mpg computer.When you get to 4th gear shift into D and let the car modify the OD.
aintain the car properly.O2 sensors at 100k miles,plugs at proper intervals,Bump up your tie pressure 4 lbs on front and rear and keep them inflated properly.Keep a clean oem air filter in the air box.And drive like you have an egg under the gas pedal. And get a G-force meter and learn to use it.
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rpm(the answer to your other question)
The extra 200 lbs for your 4 matic is costing you a few mpg,so next time unless you really need 4 WD don't get a car with it installed.:thumbsup: