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2004 W220 (S430) ; 2001 R129 (SL500) Sport
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On the S430 I own, it is the evaporator or condenser (I can't remember which) for the AC system. Look to see where the hoses run. If they run to the AC compressor, that's it. If they run to the engine or transmission, it's a cooler for that.
 
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